Google Scholar Profile. Please feel free to email me for reprints. *student. Updated September 2020.

Friesen CR, AF Kahrl, M Olsson. 2020. Sperm competition in squamate reptiles. Special Issue 50 years of sperm competition in  Philosophical Transactions of the Royal Society B


Friesen CR, DWA Noble, M Olsson. 2020. The role of oxidative stress in postcopulatory selection. Special Issue 50 years of sperm competition in  Philosophical Transactions of the Royal Society B

Friesen CR, N Rollings*, M Wilson, CM Whittington, R Shine, M Olsson. 2020. Covariation in superoxide, sperm telomere length and velocity in a polymorphic reptile. Behavioral Ecology and Sociobiology.

Lindsay WR, CR Friesen, C Sihlbom, J Bergström, E Berger, M Wilson, M Olsson. 2020. Vitellogenin offsets oxidative costs of reproduction in female painted dragon lizards. Journal of Experimental Biology.

Whittington CM, CR Friesen. 2020. The evolution of male viviparity from egg-brooding in syngnathid fishes. Biological Reviews. 


Rollings N*, HL Waye, RW Krohmer, EJ Uhrig, RT Mason, M Olsson, CM Whittington, CR Friesen. 2020. Sperm telomere length correlates with blood telomeres and body size in red-sided garter snakes, Thamnophis sirtalis parietalis. Journal of Zoology

Friesen CR, R Shine. 2019. At the invasion front, male cane toads (Rhinella marina) have smaller testes. Biology Letters.

Friesen CR, SP de Graaf, and M Olsson. 2019. The relationship of body condition, superoxide dismutase and superoxide with sperm performance. Behavioral ecology.

Rollings N*, CR Friesen, CM Whittington, R Johansson*, R Shine, M Olsson. 2019. Sex-and tissue-specific based differences in telomere length in a reptile. Ecology and Evolution.

Pauliny, A, E Miller, N Rollings*, E Wapstra, D Blomqvist, CR Friesen, M Olsson. 2018. Effects of male telomeres on probability of paternity in sand lizards. Biology Letters.

Olsson M, CR Friesen, N Rollings*, J Sudyka*, W Lindsay, CM Whittington, M Wilson. 2018. Long term effects of superoxide and DNA repair on lizard telomeres. Molecular Ecology.

Olsson M, E Wapstra, CR Friesen. 2018. Ectothermic Telomeres: it’s time they came in from the cold (Invited review). Special Issue on telomere biology in  Philosophical Transactions of the Royal Society B. 

Chen W, L Peng, L Jiang, DA Pike, CR Friesen, G Brown. 2018. High altitude frogs (Rand kukonoris) adopt a diversified bet-hedging strategy in the face of environmental unpredictability. Asian Herpetological Research 

Hurley* LL, CS McDiarmid*, CR Friesen, SC Griffith, M Rowe. 2018. Experimental heatwaves negatively impact sperm quality in the zebra finch. Proceedings of the Royal Society B: Biological Sciences. 

Olsson M, E Wapstra, CR Friesen. 2018Evolutionary Ecology of Telomeres: a Review. Annals of the New York Academy of Sciences.

Friesen CR, EJ Uhrig, E Bentz*, L Blakemore*, RT Mason. 2017. Correlated evolution of sexually selected traits:  Interspecific variation in ejaculates, sperm morphology, copulatory mate-guarding and body size in two sympatric species of garter snakes.  Behavioral Ecology and Sociobiology.

Friesen CR, Johansson, R*, Olsson M. 2017. Morph-specific metabolic rate and the timing of reproductive senescence in a color polymorphic dragon. Journal of Experimental Zoology Part A:  Ecological and Integrative Physiology. 

Friesen, CR, DR Powers, RT Mason. 2017. Using whole-group metabolic rate and behaviour to assess the energetics of courtship in red-sided garter snakes. Animal Behaviour.

McDiarmid C*, CR Friesen, C Ballen, M Olsson. 2017. Sexual colouration and sperm performance in the Australian painted dragon lizard, Ctenophorus pictus. Journal of Evolutionary Biology.

Rollings N*, EJ Uhrig, RW Krohmer, HL Waye, RT Mason, M Olsson, CMWhittington, CR Friesen. 2017. Age-related sex differences in body condition and telomere dynamics of red-sided garter snakes.  Proceedings of the Royal Society B: Biological Sciences.

Friesen CR, MR Wilson, N Rollings*, J Sudyka*, CM Whittington, M Giraudeau, M Olsson. 2017. Conditional handicaps in exuberant lizards: Bright color in aggressive males is correlated with high levels of free radicals. Frontiers in Ecology and Evolution (Invited chapter)

Rollings, N*, CR Friesen, J Sudyka*, CM Whittington, M Giraudeau, M Olsson. 2017. Telomere dynamics in a lizard with morph-specific reproductive investment and self-maintenance. Ecology and Evolution.

Friesen CR, EJ Uhrig, RT Mason, PLR Brennan. 2016. Female behaviour and the interaction of male and female genital traits mediate sperm transfer during mating. Journal of Evolutionary Biology.

Friesen CR, M Olsson. 2016. Polyandry and female post-copulatory choice. (Invited book chapter) Encyclopedia of Evolutionary Biology, Academic Press: Oxford, p307-316

M Giraudeau, CR Friesen, J Sudyka*, N Rollings*, C Whittington, M Wilson, M Olsson. 2016Aging and the cost of maintaining coloration in the Australian Painted dragon. Biology Letters.

Friesen CR, DR Powers, PE Copenhaver*, RT Mason. 2015Size dependence in non-sperm ejaculate production is reflected in daily energy expenditure and resting metabolic rate. Journal of Experimental Biology.


Friesen CR, EJ Uhrig, RT Mason. 2014Females remate more frequently when mated with sperm‐deficient males. Journal of Experimental Zoology Part A: Ecological Genetics and Physiology.

Friesen CR, AR Kerns*, RT Mason. 2014Factors influencing paternity in multiply mated female red-sided garter snakes and the persistent use of sperm stored over winter. Behavioral Ecology and Sociobiology.

Friesen CR, MK Squire*, RT Mason. 2014Intrapopulational variation of ejaculate traits and sperm depletion in red‐sided garter snakes. Journal of Zoology.

Friesen CR, EJ Uhrig, MK Squire*, RT Mason, PLR Brennan. 2014Sexual conflict over mating in red-sided garter snakes (Thamnophis sirtalis) as indicated by experimental manipulation of genitalia. Proceedings of the Royal Society of London B: Biological Sciences.


Friesen CR, RT Mason, SJ Arnold, S Estes. 2013 Patterns of sperm use in two populations of Red-sided Garter Snake (Thamnophis sirtalis parietalis) with long-term female sperm storage. Canadian Journal of Zoology.

Friesen CR, R Shine, RW Krohmer, RT Mason. 2013Not just a chastity belt: the functional significance of mating plugs in garter snakes, revisited. Biological Journal of the Linnean Society.